WWW.DISSERS.RU


...
    !

Pages:     | 1 | 2 || 4 | 5 |   ...   | 6 |

3.10. Beea ccea. B oae pee oca pocoeco aao ( pexepe peocpy) ooo cpoe eex opao opo. O pecae oo apo pecx opoo, o oopx oxo eee aa pao op poeoc. Hep opo o e y: o c ee opaa ( epo paoe) ooya. Paep cooc opaa eex opao co ca c paepa cax opo. Cae poce ep e opo c opo o pe aao xapaep cao aeo opo P. ovalis. Too y o opo ep e yacy ee oox poyo. a cee oyoc eeoo aaa y pyx opo oppepye c paepa ea (cae e coooye ep xapaep cax pyx opo Phoronopsis californica).

opa paep eex opoo opeeec e, o epo epeca epe oc ooe poy. Ta, y ox o paac e opo a ao ep. Oa x, oopa peyeceo caa c eee eeo, pyee pyo, oopa caa c eee cepe.

Ha epe oapyeo, o y paeoox o oe e eco ooo op cpoe opoo epe: y cao o aeo pyee, e y cao. Heoo ooeco ocoeoc epe opo ec ae eao-ex eo opoe (pc. 3) e ocxoeo oea ep. Hae pecx opoo oppepye y opo c xopoo pao coo epepoao poeoco cceo. papa ocyecec a cee poeocx cocyo, ocoeo a, e o oe opoa eoec ee ooa (pc. 5-I). Moa eo e ocxoeo cxoeo oe eaep caa c e, o cxoee oeo yopye a cpeopa ac, a ocxoee a opa peacop.

3.11. Hapyoe pepoye opa aeoee. Bepe a ypacpyypo ypoe ocao pae eo aoepoeao a ecx oox eo. opo e oaa oope oo eea. oaao, o coo oox eo opo c e eoeco c, opaye ce poeocx cocyo ae yoo yace ea. Ha pax aax pa ooe e ocac cocae cocyco ce oya ae poeocx cocyo. C paac oo ea eco accopoaa aoepoeaa a, e oopo pocxo a e o eoecx eo cocyco ce.

Baoepoeaa a ocae acaoo pa epoy epeca. o-oy, aoepoeaa a aaae, a ae cooae aee eeca, eoxoe paaxc oox eo.

Bopy paaxc ecx oox eo oppyec oy eo aoepoeao a (pc. 6), oop, oao, e oe, poeco y. ece ooe e opo coxpa pa ceec aapa, a ca ooee oe oece ayoceecoy y (pc. 6). Oooopee y opo ocyecec ocpeco epeoca cepaoopo. o po c oa cpoe cepaooo, oope peca coo V-opae e, e oo eo po oxop, a pyoe y. Apocoa oeaec ye V-opao e.

Moa poecco oooope, a a e aae poo y eoopx o po pa apyx pepoyx opao opo. B oae epe pee oca pocoeco aao ypacpyyp apyx pepoyx opao opo, cppoae pexep peocpy.

epapox o ec aeae ee, oope peca coo peya papaca e a ooopao oyoc apoao copo yae ypeeo pa.

Cepe aeax ee oeceae ceae poo ppeee x yaa ooopa. Opae ac aeax ee yopy a ooopae opa, yacye oppoa cepaoopo. Haeae ee ocycy y cao paeoox o, oope paca oooopee ee oy o. cao paeoox o eppopoaoy pa oepac ooopae opa, ee cooe cpoee cocoe ace, oope oy ooopoa c opao aopa ac aeax ee. Cooe cpoee ooopax opao y paeoox o oppepye c oee coo opaae cepaoopo, ex cpa apyc oaax cocooc ap oe o.

ABA 4. MPOHAHOE PABTE 4.3. Oooopee epec. Oooopee y opo ocyecec ye epea cepaoopo. Aopy yaoc oapy cepaoo yoo eoe cao Ph. harmeri, o oop o oooope ooo ee o epeca. Hepec y opo pocxo ye oooope a, a oa poa a cax poe. co paep ac o paepo ea ooo a pa.

4.4. acca o pa. opo oo e p a pa. (1) ooeaecoe pae xapaepo pyx opo, oax oe oo pya. co eaex epec ca, paep o 90 , epa aaa a acya. (2) Pae c aae xapaepo eox opo, yx oe epoo pya. co poypyex co, aep 100 , epa aaa a aopoxa. (3) eopooe pae xapaepo oo ooo a Phoronis ovalis. co e oee 20, aep 125 . a e aae, e aec.

Pc. 6. Cxe oceoaex ca ooeea Phoronopsis harmeri peopaoa opyax eo aoepoeao a.

poo pacpocpaeoe epaype ee o cpao e poe y opo pao oo. Oo ocoao a peyae pao Paeap (Rattenbury, 1954), oeo a Phoronopsis viridis (a co Phoronopsis harmeri). B peyae epecceoa pa oo e a oyeao oaao, o poee a poeae a paaoe (pc. 7). o aee oepeo a e c py o pa P. ijimai.

Oapyeo, o eoope ocoeoc poe oppepy c o pa (ep op). p ooeaeco e pa opo eepoo ee poe poxo o eoop yo aao-eeao oc apoa, o ooe coppoa ooc yp poa ye a ca 16 acoepo (pc. 7E).

p pa c aae opo eepoo oo ee poe poxo cpoo epoao apae, peyae eo apo popeae opy yxcoo ac. Taa opa pa coxpaec a oee ox cax oeceae oay yaoy apoe poax coex.

Ha epe oapyeo, o y opo eoepa aaaec yx aao: epeeo aeo. epe eoepa aao oppyec a ca acpy a ce cee eo epee ce apxeepoa (pc. 7M, H). e ocec ay eoecy cy pepoooo yaeoo eoo. a eoepa aao oppyec a ca ooo .

O oec a eapoe opcaoe ae ce cpee , o ec opayec epoe ye. aao pace, oypoaec o ce , oae e c oo, popea oooopay opy. oceeo pa oo cac cac a epao copoe ea , e oppyec ece eap eeep. a aao eoep ae cy yooo eoa.

a coa eoep (epe a) xapaep ox pyx py Bilateria. Ta, y Spiralia a eoepa pocxo apaa D, oee o e 4d, a a epe co eoep oo paoa e, pocxoe o acoepo 2a, 2b, 2c 3a, 3b, ae aao ycyaype o (Lillie, 1895; Biggelaar, 1983; Boyer et al., 1998; Henry, Martindale, 1998; Lartillot et al., 2002a). xopox a eoepa o eoepa yoo-xocoo o, a epe o a aaea pexopaa eoepa (Adelman, 1922; Seifert et al., 1993; Kiecker, Niehrs, 2001). a coa eoep xapaep paoopax, e ocoa ac eoep pocxo ae oac apoeoo ca (c. Manton, 1928, 1934; Nair, 1941, 1956; Weygoldt, 1960, 1961; Anderson, 1967;

Behesch, 1969). B o e pe epee ac apoa y ox op oeeo oocoee a aaeo peaeao eoep (c.

Manton, 1934; Nair, 1939, 1941, 1949, 1956; Weygoldt, 1960, 1961; Benesch, 1969), oopa ae ycyaypy epxe y, eoa, a ae ax ceeo. B o c epeco oe, o oac epeeo aeo eoepax aao y ooox ecooox oeea cpecc oeoocx eo oooo Brachiury, goosecoid fork head (Weigel et al., 1989; Artinger et al., 1997; Filosa et al., 1997; Bassham, Postlethwait, 2000; Tagawa et al., 1998, Pc. 7. Pae Phoronopsis harmeri, oopa c x poo.

2001; Kusch, Reuter, 1999; Technau, 2001; Lartillot et al., 2002b; Takade et al., 2002). Bepoo, epe a eoepae aa Bilateria, cae c epe a oa eeoo acoopa, o poo ye pacee oeopaoo epa aa eoep paao-cepoo pea.

Mooa aopoxa aopoa (a opo) ee xapaepy -opay opy (pc. 7), y ee eec coo e, epeoa a oe, pepooo ( o poa Phoronopsis) yo eo, pepooo ocepooo pece yp pooep. ocee aaac y ooo a ape a oep c epo-aepax copo o ayca. o a oee ox cax oec eopox () a oooop opa (pc. 8). peopa yp o opo oe ooopoa c poopoxo o Spiralia, a ocopa yp c eapoxo. B o e pe peopa ocopa pece yp aopox oy ooopoa c cooecy pec ypa o opopox.

Pc. 8. opo c ooeaec pae (A) pae c aae (). oopa x ox.

Ha epe yaoc eo poce copypoa oppe ey o pa o y opo. o c ooeaec pae (pc. 8A) e ooceo pye paep (oee 1 ), popae opo, op acoe, ooe co yae (oee 24) oo (2.53 ec.) axoc aoe.

o, oopx xapaepo aae poo poe yae (pc.8), a ea o ee 1 . e oe, epopae (cce) epoae opo ea, acoe y, co yae e peae 14. Moo pa eopox cy a oooop opao. axoc aoe e oee 1 eca.

eopoa oaa a xapaepa ooo a opo Phoronis ovalis, o cao c yeee paepo ea pocoo ooo , a cece, eoooc poypoa ooe oeco .

ABA 5. MOPOO, AHATOM, TPACTPTPA HO OPOH 5.1. Mopoo o. peeo ocae ee opoo o opo, oocxc 6 a (Phoronis ijimai, P. psammophila, P. muelleri, Phoronopsis harmeri, Ph. albomaculata, Ph. californica), a a e o, oa paeoc e a ycaoea (Actinotrocha sp., aco aopox o-acoo op aopox aoa epoo op). oca opo ycpoe o oey ay. Ba opeee o opo cac ceye ocoeoc ee opoo: a ea, ocaea peeaopooy epoy, popaoc / epopaoc opoo, ae oaa ea opoo, acaoe co yae, ae / ocyce aao ex yae. a ccea ocoeoc opaa o oocc: opa pepoooo eoa, co eeox epy eya (o a), co coe ppoo, ae / ocyce poaoo opaa.

peeo ocae aco aopox, oapyeo oaco ope. poe eoo pyx paepo (3.5 ) a ea eoope ocoeoc aaoeco opaa, cocee poc o. B o ce y ee oapye aa oa. a aa eoea acx eaecx o (oope ec eooocx, pecx epe, cyy, oyxopox eoopx ooox), oop, o aey e, ca e c cyecoae acx pocx op, a co coe popae eaopoa.

5.2. epc. peeo ocae ooo cpoe epca Actinotrocha harmeri, A. vancouverensis, A. sp. ooe cceoa oe a ax opo epe. B ee oo e ecoo yaco epca, oope paac ocoeoc oo opaa: peopaa oac, opaoe oe, yaa, yoe, epc eopoxa, eopox, oooaaoe oo. opo e o opo opaoa ooap ea (pc. 9A). oepa peopao oac, opaoo o oac eopoxa opaoaa ooap eao-e ea. Moae poxo aax acx eo, a a e oppy pooe (pc. 9A).

peopa, ocopa pece yp eopox a e opaoa ooap ea, o cae aopox o opopox ox. B epce aopox ooce ycee e, oope e cxooe cpoee c o ea ea oac o x ae opoa opy ya ae acoa.

Pages:     | 1 | 2 || 4 | 5 |   ...   | 6 |



2011 www.dissers.ru -

, .
, , , , 1-2 .