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a npaax pyonucu Teepea Eea Hoaea opo (Phoronida):

cpoee, pae, poa aya, oe 03.00.08 ooo Aopeepa ccepa a cocae yeo cee oopa ooecx ay Moca 2008 Paoa oea a aepe ooo ecooox ooecoo ayea Mococoo ocyapceoo yepcea e M.B. ooocoa.

Hay ocya oop ooecx ay, poeccop, e-opp. PAH Maaxo Bap Bace Oae ooe: oop ooecx ay, aae PAH Apao Ape Bapo, cy oo op BO PAH;

oop ooecx ay, poeccop Maae p Bopo, oooec cy PAH;

oop ooecx ay pae Tp Aao, cy poe oo o . A.H. Ceepoa PAH Beya opaa: Ca-eepypc ocyapce yepce.

aa cococ 27 op 2008 . 15 . 30 . a acea ccepaooo coea 501.001.20 Mococo ocyapceo yepcee e M.B. ooocoa o apecy:

119991, Moca, M e M.B. ooocoa, ooec aye.

C ccepae oo oaoc oee ooecoo ayea M e M.B. ooocoa.

Aopeepa paoca 26 cep 2008 .

e cepeap ccepaooo coea aa ooecx ay apcoa apca aoa Ayaoc e. opo opce opa, ee ceceoe pacpocpaee oapyee o cex aaopx Mpooo oeaa a cee aapecx po. Phoronida ooo apca, cocae oopoo acaec ceo 12 o.

Heoope x o oy oca opoo ceoc o 94000 ./. (Emig, 1982). a oaa peya ax cceoa, opx Pocc oae 8 o opo, oope opay ope ocee aaopx epoo ocoo ope (Capao p., 1967; Tapaco, 1978). -a oo ceoc opo opy eoopx opcx cooecax (ocoeo a x pyax) opee x coca. oco opo ee aopox o, oope y aoe 13 eca. a o pe o oca oo ceoc coca aey o epoaoa. Mooceoc opo x o eae y pyy ao p ye cocaa ox aox cooec, yee opaoopa poee pax pyx poooecx, ooecx oooecx pao.

opo e eo a cpoe, a caa pya ceo epeca c o pe cpaeo aao poo. o cooyoc pao oa aae poeyooe ooee ey epopo (Protostomia) opopo (Deuterostomia) o. B c c ooee opo ccee ooo apca ooe pe ocaaoc ece. oco cceoaee ca opo opopox ox a ocoa opaa eoeco cce ocoeoce poo. B eoopx pyx oooecx pyoocax opo paccapac a oa aoee px py Bilateria, cao paccopee aepao cepx ox aaec c opo (Remane et al., 1975). Haoo aco Lophophorata, ya ece c opoa a ao paxoo, ooe pe paccapac paccapaec eoop cceoae o cx op a aa cpe cex Radiata (Jefferies, 1986; Ax 1989, 2000). Oao, coaco o a, oye copee eoa oeypo oee, opo paccapac a pya, pocea epopo o aea, oca, oc ep (Halanych et al., 1995). a oa pe oa e aa ax oepe co copo acceco aao poo, ooy ooee opo ccee ooo apca a ocaec ece, a pap pecaex opooo oeypx oeeo e oo e yeaec, a yeaec co peee.

e aa cceoa. e acoe pao ec peocpy opoo, pa, acooeco cpyyp, aa cpoe oe Phoronida oeoo a ooo apca.

B aa cceoa xoo:

1. ocyec oy peocpy aaoeco opaa pocx opo;

2. y ypacpyypy cex ae opao pocx ox;

3. cceoa poaoe pae opo;

4. y aaoecy opaa ypacpyypy cex ae opao y o opo, paeax pa poa;

5. poec yee eaopoa o opo poce cyy eaopoe oex opao;

6. coca cocc poo ay pocx opo c opo oca opooecx aaoecx ocoeoce ox pax aaop Mpooo oeaa. Coca pacpee ooee oca o x opeee;

7. y ooe pae opo, a a e opoo aao aopox aox po, copax xoe ce cyo PAH, coca oe oca cex ecx acoey pee o opo x opeee;

8. a ocee eooy ay cpoe opo epecope oeeece ooe o py c yeo ax opoo oeypo oee.

Haya oa. peoea opaa cxea oce aa cpoe opo oa oe x oe, oopa oee paoe cpaex aaoo oe oeype ae.

B xoe eax cceoa opoo, pocoeco aao ypacpyyp opo oapyeo, o pya eo xapaepyec peao po opooeco, cooeco ooeco opaae. Bce a cce opao eocppy apxaoe cpoee: opo epc opaoa ooap ea, oopx aa eo ya ec oa epoe; ce ee epo cce aea oe opooo e e opaa paepaoo ecyca; oepa e eya cpee opaoa eao-e ooap ea, oooe cpoee oep ca opo ocooe ooee cpe cex Bilateria.

Cpoee ooo cce opo a e eocppye ep apxaoc: ocoo cyecyx oa y opo e, ooe e paac eo eoeco c, opayx ce poeocx cocyo.

Bepe poee opo-yoa aa eoaeoo aapaa opo e exa paepo cepea ac.

Bepe oapye copypoa eoope aooepoc poaoo pa opo, oaa paa poe a. Bepe oapyeo, o opo eepoo oo ee poe poxo o-paoy acoc o a pa.

Bepe oapyeo a coa a cocoa oppoa eoeco eoep. Bepe eeo a a aopox o opo, paaxc ocoeoc opoo, aao oo. Bea ope ey o pa o .

Bepe poeeoe eaoe cceoae cex cce opao o opo ooo p yax ep x opaa: ooape eao-ee e opoax ( o ce pecx ypax) eoecx cax; ae pao poeoco cce (o eoa epa cpoe aox o); ae yx o opaa eoeco cce, o ce pepo cce y o poa Phoronopsis.

Bepe ye eaopo Phoronopsis harmeri. Bepe opo oaao ooee ex yae pocax oo ox yae. Bepe poceea peeceoc ey eapeo cceo pocx opo x o.

Bepe poceea cya eaopoe eex opao opo peoea cxea x pa oeee. Bepe ocoepo ycaoeo, o aepae eeep eaopoe oc peyae ao poepa eo eoeco c yooo eoa. Bepe peoea opaa cxea oeeecoo pa aepax eeepe aa cpoe opo.

Haeo ocao 2 ox ay a opo ( ooee 10 paee ec). Bepe ae oca opo apeco accee Mpooo oeaa, epoo ope, y xooeacoo oepe aa. 3 a opo epe apecppoa opx Pocc ( ooee 5 ec). Bepe cocae o opeee cex ecx o opo c yeo ocoeoce aaoeco opaa opo pax aaop Mpooo oeaa.

Bepe ocaa opoo aao o opo, ocoo oapyaex aoe epoo op a. Tepe Oxocoo op. Oapye paee eece , peooeo paeae a Phoronopsis albomaculata Ph. californica.

Bepe opo oca eoe acx o poaapoaa poea aa o opcx ox. Bepe opoo oca oceoaee ca ooo pa pex aoee accox o opo ocoo op. Bepe cocae opeee o opo. Bepe aece poooo opeeeoo paa peoeo cooa opy pepoooo eoa.

Ha ay occ ceye ooe.

Hoe pecae o ae cpoe, opooeco, cooeco ooeco opaa opo.

Hoe pecae o e poaoo pa opo.

o cocc poo ay opo c ocae cex o e opeee pocx o.

Hoe pecae o ae cpoe Lophophorata a ooepx (a e apxepx) ox x ooe cpe Lophotrochozoa Apoa pao ya. Peya pao ooe a II Bcepoccco coye ooece poe ooeea, Moca, 2000 .; a oepe eoopco ooeco ca 2000 .; a ayx ooyax cya oo op BO PAH 19992005 .; a eo oepe a A.B. aoa poe ooo opoo ox H PAH; a 7- Meyapoo coye Larval Biology 2006 . . Coos Bay (Opeo, CA); a ex a B.. Baa Mopca ooo apaoo: oc op aaco ocyapceo yepcee 2007 .; a aceax aep ooo ecooox ooecoo ayea M e M.B. ooocoa.

o ee ccepa oyoa 32 pao ( x 28 cae peepyex ypaax).

Cpyypa oe pao. ccepa pecae pyoc 2-x oax. ep o coep ec ocoo ac poe, opo cpa ocoo ac poe. Tec ocoo ac coco ee, 7 a, oo cca epayp.

Cco epayp ae 401 aae, oopx 342 a ocpax ax. To ep coep 514 cp., o opo 287 cp.

aoapoc. Aop paae yoy aoapoc ayoy ocyay yooyaaeoy Maaxoy Bapy Bacey, a ae ooce oea py, oaa oo oye ayoo aepaa, epayp, ex coeo, xopoeo acpoe oeoo acpo: y B.B., epey A.B., Paaecoy B.., Apaoy A.B., Mxaoy .., Pycy ., peo O., Macaoo C., o M., Coooy C., epay ., Cooey ., P. epy, C. Caaae, . y p. Aop yoo paee copya aep ooo ecooox ooecoo ayea M e M.B. ooocoa aopaop poo cya oo op BO PAH. Ocoy aoapoc aop paae.H. aoy, A.. oaoy ce copya eaepao aopaop epoo poco M e M.B. ooocoa.

ABA 1. CTOP EH OPOH B ae opoo paccope ocoe a ye cpoe, pa acoo opo.

ABA 2. MATEPA METO Maepao pao ocy poce opo 8 o pax paoo Mpooo oeaa: Phoronis ovalis, P. ijimai, P. hippocrepia, P. psammophila, P. australis, P. svetlanae, Phoronopsis harmeri, Ph.

malakhovi, copae aopo o, a ae oopae oex cyo Poccco aae ay. Bce opo pax ec oa ye p oo cooecx eoo (ceo oyeo ooo 140 cep oepex, 3 cep caax 2 cep poax cpeo).

Toa opoo pax yaco ea py cceoaa a pepe P. ijimai, P. hippocrepia, Ph. harmeri eoa capye epoo poco (CM). Tooe cpoee cex cce opao cceoao eoa pacccoo epoo poco (TM) a pepe Ph. harmeri. Pae yeo a pepe yx o P. ijimai Ph. harmeri.

Maepao pao ae ocy 9 o opo, paee oooopex oopae aox po, x pax aaop. Bce cceoa cooec eoa (ceo oyeo 15 caax, 20 oepex 6 poax cep cpeo). Toa opoo eoa CM yea y o 5 o. Tooe cpoee cex cce opao yeo a A. vancouverensis, A. sp., A. harmeri. coxec eoa ( e oe aao eoo aapaa) c peee aepo ooao poco cceoa P.

ijimai Ph. harmeri. Meaopo opo ye a pepe Ph. harmeri.

ABA 3. MOPOO, AHATOM, TPACTPTPA BPOCX OPOH 3.1. Tpya opoo oo ea. opo opay e ooece py: oae o cycpae (pc. 1A) cepe opo, oae ax paoax oco (pc.

1). Tpy opo coco pex coe (pc. 1B) opaoa c yoocaxapa, oco cyoyoocaxapa ea. Bepe oaao, o co py coco ox cycoe (cc) (pc. 1B), a oopx opaoa cea epeeee ox p. Tooe cpoee pyo opo aeo poe, e y pyx yox ox (apep, oxe eceep). Tpy opo e e oecy, a pecy opy, ae cea pacpe a ae oe. Moo peoo, o poecce poca opo coco aco pacop aoo e aepa aeo yaca py. Paep ea ox appy o 1 o 50 c y. Moe eo ox oe opaeeo a yac, paaec ocoeoc opoo: epe (), ecy ooop co yaa (), epe yo (y), a yo (y) ayy (a) (pc. 1).

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